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3 Translational decision framework for VLP vaccine engineering The four evaluative dimensions introduced in Sections 2.1 Extra-VLP protein antigen display technologies and 2.2 Intra-VLP nucleic acid encapsulation technologiescargo flexibility, loading specificity, functional efficiency, and manufacturabilityconverge into a translational design logic integrating antigen display and nucleic acid encapsulation within a unified development pathway for rational VLP vaccine engineering (Figure 4)