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In zebrafish, Grx2 is an essential protein for successful embryogenesis, regulating neuronal survival and axon growth [92] as well as angiogenesis through S-glutathionylation of the histone-deacetylase sirtuin1 [93,94]

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Thus phosphorylation of TDP-43, in addition to cleavage, may be additional mechanisms to regulate TDP-43 proteostasis in the event of SUMOylation-pathway failure

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